The tomato (Solanum lycopersicum) was engineered to have a purple coloration and contain increased levels of antioxidants. To achieve this, the tomato expresses the Antirrhinum majus transcription factors ROSEA1 and DELILA, which interact to upregulate the the genes associated with anthocyanin biosynthesis in the tomato. As such, the levels of  tomato anthocyanins delphinidin 3-O-(coumaroyl) rutinoside-5-O-glucoside and petunidin 3-O-(coumaroyl) rutinoside-5-O-glucoside are elevated in the flesh of the modified tomato fruit, resulting in a deep purple colour. In addition to the transcription factors, the modified tomato also contains an Escherichia coli neomycin phosphotransferase II gene cassette, which was used for neomycin selection during transformation.